Reef Sharks and Rays, Part 3 Codexery

Himantura fluviatilis

An obscure, Endangered stingray of the Ganges River system.

Himantura fluviatilis

The Ganges whip ray, *Himantura fluviatilis*, is a poorly understood stingray from the family Dasyatidae. It appears to live only in the Ganges River system and nearby coastal waters. Its identity is uncertain: it looks very much like the giant freshwater stingray of Southeast Asia (*Himantura chaophraya*), and the two may actually be the same species. The ray has an oval pectoral disc and a long, pointed snout. It grows to about 1.4 meters (4.6 feet) across. Its upper side is a plain dark color, while the underside is lighter with wide, dark bands along the edges of the disc. The International Union for Conservation of Nature (IUCN) considers it Endangered because it faces overfishing and habitat loss in its densely populated range.

**Taxonomy** In 1822, Scottish naturalist Francis Buchanan-Hamilton wrote about a freshwater stingray from the Ganges that he called *Raia fluviatilis*, but he never formally described it because he lacked an illustration. He only noted that it resembled *Raia lymma* (now *Taeniura lymma*) and behaved like *Raia aquila* (now *Myliobatis aquila*). Later researchers referred to it as *Trygon fluviatilis* and then *Himantura fluviatilis*, though many have doubted the name’s validity due to Hamilton’s sparse description. The IUCN Red List still treats *H. fluviatilis* as a synonym of the cowtail stingray. It belongs to the same species group as *H. chaophraya*, and the two might be synonymous, but no specimens exist to compare.

**Distribution and habitat** Hamilton reported that the Ganges stingray was common in the Ganges system from the estuary up to the Kanpur area, about 1,600 kilometers (1,000 miles) inland from the tidal limit. Later records also place it in shallow bays of the Bay of Bengal and at depths of 37–55 meters (120–180 feet) near Madras. This ray is probably completely cut off from other freshwater stingray populations in Southeast Asia.

**Description** The Ganges stingray has a very thin, oval pectoral disc that is slightly wider than long, with broadly rounded edges. Its snout tapers to a narrow point and extends well beyond the disc, measuring more than twice the distance between its very small eyes. The whip-like tail lacks fin folds and is less than twice the disc length. The entire upper surface, except the pelvic fins, is covered with dermal denticles, which become larger on the head.

Quick Facts

Genus
Himantura
Species
fluviatilis

Facts from the source article.

Lore & Background

Scottish physician, geographer, and naturalist Francis Buchanan-Hamilton, in his 1822 account of Ganges River fishes, made reference to a species of freshwater stingray he named Raia fluviatilis. However, he deferred making a description until an illustration could be made, an opportunity that never arose, and as a result his account only notes that the ray 'has a strong resemblance to Raia lymma [=Taeniura lymma] [and] has nearly the same manners as Raia aquila [=Myliobatis aquila]'. Subsequent authors have published accounts of this species under the name Trygon fluviatilis, and then Himantura fluviatilis, while noting that the name has questionable validity due to the paucity of Hamilton's description. The IUCN Red List still lists H. fluviatilis as a synonym of the Cowtail stingray. The Ganges stingray belongs to the same species group as H. chaophraya; the two species may in fact be synonymous, but this cannot yet be confirmed as no specimens are available for comparison.

The Ganges stingray has a very thin, oval pectoral fin disc slightly wider than long, with broadly rounded lateral margins. The snout tapers to a narrow point and projects considerably beyond the disc, measuring over twice as long as the distance between the eyes, which are very small. The tail is whip-like, lacks fin folds, and is less than twice as long as the disc. The entire upper surface, except for the pelvic fins, is covered by dermal denticles that become larger on the head. Some of the denticles on the posterior half of the body are also enlarged, and bear sharp spines. Large areas of the underside are covered by minute denticles. The coloration is a plain purplish grey above, becoming darker towards the disc margins, and light below with broad, irregular dark bands bordering the lateral margins. This is a large species, reaching a width of at least 1.4 m (4.6 ft).

Hamilton noted that the Ganges stingray was common in the Ganges River system from the estuary to the area around Kanpur, some 1,600 km (1,000 mi) from the furthest extent of the tide. Later authors have also reported it from shallow bays in Bay of Bengal, and at a depth of 37–55 m (120–180 ft) near Madras. This ray is likely completely isolated from other populations of freshwater stingrays in various parts of Southeast Asia.

Reader's Guide

The Ganges stingray's significance lies in its extreme obscurity and the taxonomic uncertainty that has surrounded it since its first mention in 1822. Only a few specimens are recorded in literature, none of which remain to the present day. It has often been mistaken as a disjunct population of Urogymnus polylepis or even the Cowtail stingray. The International Union for Conservation of Nature (IUCN) has assessed the Ganges stingray as Endangered: it has a restricted distribution in a heavily populated region, which renders it susceptible to overfishing and habitat degradation. Hamilton noted that this ray was often sold at Kanpur markets, and that its tail spine can inflict 'very dangerous wounds'. It probably continues to be caught as bycatch within its range. The Ganges river stingray is a protected species in India, being listed on Schedule I of the Indian Wildlife (Protection) Act of 1972, a status that it shares with the rare Ganges shark. A reported sighting of a dead Ganges River stingray in 2002 states that it had a 76 cm (30 in) disc width, found dead in the Chambal River. A 'Giant stingray' was caught in 2019 in a river in Odisha, India; this species of Indian Giant freshwater stingray species are rarely found because most of the rivers which they live are severely polluted due to human interaction.

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