Psocoptera
Primitive paraneopterans, including booklice and barklice.
Psocoptera are a paraphyletic group of insects commonly known as booklice, barklice, or barkflies. They are often regarded as the most primitive of the paraneopterans, with more than 5,500 species in 41 families across three suborders. Their name originates from Greek words meaning 'gnawed' and 'wings.' In modern systematics, Psocoptera has been replaced with Psocodea, as the parasitic lice (Phthiraptera) evolved from within the psocopteran suborder Troctomorpha, making Psocoptera paraphyletic.
- Common name
- Booklice, barklice, barkflies
- Number of species
- More than 5,500
- Size range
- 1–10 mm (1/32–13/32 in)
- Suborders
- Three
- Families
- 41
- Known for
- Feeding on fungi, algae, lichen, and starchy materials; some species are pests of stored products
Lore & Background
Psocoptera are small, scavenging insects with a generalized body plan. They have chewing mandibles, a modified maxillary rod used to brace while scraping detritus, a swollen forehead, large compound eyes, three ocelli, and a soft segmented abdomen. Some species spin silk from mouth glands. Their legs are slender and adapted for jumping, not gripping. Wings, if present, are held tent-like over the body, with simple venation. Many individuals within a species show variation, including winglessness and differences in thorax shape or setae development. Many psocids are parthenogenetic, and male presence may vary between races.
Reader's Guide
Psocoptera are significant as primitive paraneopterans and as a paraphyletic group that gave rise to the parasitic lice (Phthiraptera). Their classification was revised in the 2000s based on morphological and molecular phylogenetic evidence, leading to the inclusion of Phthiraptera within Psocodea. Ecologically, psocids are scavengers that feed on fungi, algae, lichen, and detritus, contributing to decomposition, especially in environments with few predators. The nymph of Psilopsocus mimulus is the first known wood-boring psocopteran, creating burrows that other organisms may use. While most species are woodland insects with little human contact, some, like Liposcelis bostrychophila, are pests of stored products. Booklice are commonly found among old books, feeding on paste bindings, and can infest food storage areas. Modified atmospheres during packing have been studied to control their recurrence in food manufacturing.
Did You Know?
- The name Psocoptera comes from Greek words meaning 'gnawed' and 'wings.'
- Booklice received their common name because they are commonly found among old books, feeding on the paste used in binding.
- Some psocids can spin silk from glands in their mouth and may festoon large sections of tree trunks with silk.
More in Insect Families 1-20
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