Marine Snails: Ceriths, Nassas and Conchs, Part 2 Codexery

Demoulia

A genus of marine snails in the family Nassariidae.

Demoulia

Demoulia is a genus of marine gastropod mollusks, commonly known as dog whelks or Nassa mud snails, and belongs to the family Nassariidae. The shell of these sea snails is ovate and globose, with a soft, downy outer layer called an epidermis. Its spire is short and conical, ending in a papilla-like tip, and the whorls are flattened. The opening of the shell is oval, with a thickened inner lip that has a ridge at the back, while the outer lip is narrowed, thickened on the outside, and lined with folds on the inside. Species in this genus include *Demoulia abbreviata*, *Demoulia nataliae*, *Demoulia obtusata*, *Demoulia tryoni*, *Demoulia ventricosa*, and several extinct forms such as *Demoulia conglobata*, *Demoulia pupa*, and *Demoulia zbyszewskii*. Some names once considered separate species, like *Demoulia kurodai*, *Demoulia pulchra*, and *Demoulia retusa*, are now recognized as synonyms of other species.

Family
Nassariidae
Type genus
Demoulia
Type species
Demoulia abbreviata (Gmelin, 1791)
Extant species
Demoulia abbreviata, Demoulia nataliae, Demoulia obtusata, Demoulia tryoni, Demoulia ventricosa
Fossil species
Demoulia conglobata, Demoulia pupa, Demoulia zbyszewskii

Lore & Background

The genus Demoulia was established within the Nassariidae, a family commonly known as Nassa mud snails or dog whelks. Its shells are ovate-globose and covered with a downy epidermis. The spire is short and conical, ending in a papillary apex, and the whorls are depressed. The aperture is ovate, with a thickened inner lip bearing a ridge posteriorly, and the outer lip is contracted, thickened externally, and plicated internally.

Several species have been described, both living and fossil. Among the living species are Demoulia abbreviata, Demoulia nataliae, Demoulia obtusata, Demoulia tryoni, and Demoulia ventricosa. Fossil species include Demoulia conglobata, Demoulia pupa, and Demoulia zbyszewskii. Some names have been brought into synonymy: Demoulia kurodai is a synonym of Nassarius sufflatus, Demoulia pulchra is a synonym of Demoulia obtusata, and Demoulia retusa is a synonym of Demoulia ventricosa.

Reader's Guide

Demoulia represents a distinct lineage within the Nassariidae, contributing to the understanding of mud snail diversity. The genus includes both extant and fossil species, indicating a long evolutionary history. The morphological features—such as the downy epidermis, papillary apex, and plicated outer lip—help distinguish Demoulia from related genera. The synonymies reflect taxonomic revisions that clarify species boundaries. The genus has been included in phylogenetic studies of the Nassariidae, which have revisited the family's systematics. Its fossil record, with species like Demoulia conglobata and Demoulia pupa, provides insight into past marine environments. The genus remains a subject of taxonomic interest, with ongoing refinements to its species list.

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